By H. W. Doelle
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Extra resources for Bacterial Metabolism
THERMODYNAMICS OF BIOLOGICAL REACTIONS and various lyases and synthetases. 22). The fate of this substance depends on the nature of the enzyme protein to which the pyridoxal phosphate is attached and on the group R. ^Γ^ I HC I H 2 C OOC. 22 Thiamine Pyrophosphate. 23). The structural formula of thiamine has two heterocyclic rings which are not condensed: one pyrimidine and one thiazol ring. It there fore always carries a charge. 24). 25). 24 H3C/C^N COOH *co2 Y N^ V C C \ (X H 3 C^ ^Ν^ ^NH 2 ^^^^-N C—CH3 OH OH ° \ / C .
Reductant f-^_ ADP ATP chlorophyll FIG. 3. ]. present in the chromatophores, is changed into NADPH + H + by a vigorous photosynthetic reduction of NADP + and could convert reduced NADP + to NAD + . Cytochrome c-552 could in this case take the place of ferredoxin. The presence of a pyridine nucleotide transhydrogenase has been observed in Rhodopseudomonas sphéroïdes(91), where the trapping of reduced NAD + completely inhibited the reduction of NADP + although NADP + was preferen tially used over NAD + .
The reduced form of ubiquinone (ubiquinol) is oxidized through the cytochrome system (14). The oxidation depends on cytochrome c and cytochrome oxidase and is quite often coupled to phosphorylation. Vitamin K has been reported in the electron transport processes of photosynthetic phosphorylation in bacteria, although ferredoxin may replace it in this role as will be seen later. Phosphate Carriers. The two most important types of transport reaction are electron transport and phosphate transport.